- Ecology, primary productivity and systematics of freshwater algae. Paleolimnology.
- Population and community ecology of protozoans.
- Aquatic ecology, including nutrient dynamics and effects of environmental contaminants.
- The invertebrate fauna and ecology of running water. The fate of organic matter in streams. Ecology of freshwater amphipods and stoneflies.
- Freshwater biology; ecology of Chironomidae and biology of freshwater molluscs.
- Ecology of limnetic zooplankton and benthic invertebrates.
- Biology of salmonid fishes and fluviatile forage fishes. Winter ecology of freshwater fish.
- Biotelemetry of fish and wildlife.
- Environmental toxicology: effects of toxicants on freshwater and terrestrial ecosystems and organisms. Pesticide, organic-chemical and heavy-metal effects.
- Ecological and toxicological impact of solar ultraviolet and visible radiation.
- Ecology of juvenile marine fish species.
- Migratory behaviour of diadromous fish species.
- Environmental toxicology: development of bioassay systems for quantifying lethal and sublethal damage from toxicants using algae, microorganisms and cell tissue cultures. Studies on mode of action, degradation and cycling of toxicants. Physiological responses to toxic chemicals.
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